A Cooperative Species
Model was written in NetLogo 6.0.2
•
Viewed 737 times
•
Downloaded 59 times
•
Run 0 times
Do you have questions or comments about this model? Ask them here! (You'll first need to log in.)
Comments and Questions
Please start the discussion about this model!
(You'll first need to log in.)
Click to Run Model
globals [ combo replacement_rate ; set at 5% mutation_rate ;; 2% groups ; just a list of numbers from 1 to N_group p_links ;; total possible links given n nodes p_networks ;; total possible networks from n nodes leaders ;; list of "leaders" in round, to serve as anchors for the layout-spring algorithm followers ;; everyone who isn't a leader avwithingroupvar ;; average within group variance betweenvar ;between-group variance variance_ratio ;; Fst = var(pj) / [Avvar(pij) + var(pj)] = population-wide measure of the degree of non-randomness in who interacts with whom; aka *inbreeding coefficient* ;; = differences in the probability of being paired with an altruist conditional on being an altruist, and the probability of being paired with an altruist conditional on ;; being a non-altruist (defector). ;; one expects cooperation to prevail when Fst > c / b. pop_change ;; expected change in fraction of altruists ] breed [cooperators cooperator] breed [defectors defector] undirected-link-breed [wlinks wlink] ;within-group links undirected-link-breed [blinks blink] ;between-group links ;links-own [memories] turtles-own [ earnings ;; accumulated payoffs payoff N_Neighbors mycosts mybenefits t_threshold groupid ;; groups 1 --> N_groups group_coop ;; previous number of contributors/cooperators in the previous round, withint he group; ;;should be the same for turtles of the same group sorted contrite ;; number = 0 originally, if accidentalyl makes a mistake and defects, then set to 2, which means agent will cooperate next 2 rounds automatically. wingroupvar ;; within group variance of altruism p_i ;; probabilistic interaction; likelihood thta other turtles will interact with this turtle.. test ] to setup clear-all reset-ticks set-default-shape turtles "face happy" set groups [] set leaders [] set followers [] let g 1 repeat N_groups [ set groups lput g groups set g g + 1] create-turtles (N_groups * size_n) [ while [any? other turtles-here] [ let empty_patch one-of patches with [any? turtles-here = false] move-to empty_patch ] set sorted false set groupid 0 ] setup-neighbors ask turtles [ let cnt size_n ;; (i.e. n) let t random cnt + 1 ;; i.e. between 0 and n, n = # in group. Interesting to test differences using n and n-1 as ;; used by Bowles and Gintis. When using n, turtles with t = n will only cooperate if everybody cooperated in the previous ;; round, including oneself! A turtle with t = n + 1 is a DEFECTOR, set below. set t_threshold t set contrite 0 set breed cooperators set color yellow set size 1 set group_coop size_n ;; turtles act initially as if everybody in group cooperated last round ] let pop count turtles let num_d (Percent_Defectors / 100) * pop let new_defectors n-of num_d turtles ask new_defectors [set breed defectors set shape "face sad" set size 1.5 set color red set t_threshold size_n + 1 set group_coop size_n] ;if GAME = "Pairwise Prisoners Dilemma Game" [ask turtles [create-links-with other turtles [set hidden? true]]] ;layout end to start if count turtles > 0 [ if GAME = "Public Goods Game" [Public_goods_game] if GAME = "Pairwise Prisoners Dilemma Game" [PD_pairing] if Replicator_Dynamics? = true AND count cooperators > 0 AND count defectors > 0 [replicator_dynamics] if count turtles > 0 [ if reassortment? = true [setup-neighbors] if starvation? = true [dying-turtles] if kill_defectors? = true [kill-d] ask turtles [if contrite? = true[ ; cooperate if defected in error from previous 1-2 rounds if contrite > 0 [set contrite contrite - 1]]] update-plots ;layout tick ] ] end to setup-neighbors if GAME = "Public Goods Game" [assign_groups] if GAME = "Pairwise Prisoners Dilemma Game" [ create-pairs ;layout ] end to create-pairs ask turtles[ if PD_assortment = "Random" [create-pairs-random] if PD_assortment = "Fixed" [create-pairs-fixed]] end to create-pairs-random set n_neighbors other turtles ;; This will end up being proportional to the population distribution end to create-pairs-fixed ifelse [breed] of self = cooperators [ let p Probability_of_Altruist_meeting_Altruist let r random 100 ifelse r < p [set n_neighbors other cooperators][set n_neighbors defectors]] [let p Probability_of_Defector_meeting_Altruist let r random 100 ifelse r < p [set n_neighbors cooperators] [set n_neighbors other defectors]] end to-report find-partner let partner one-of N_Neighbors if partner = nobody [set partner one-of turtles] report partner end to assign_groups ask turtles [setxy random-pxcor random-pycor while [any? other turtles-here] [ let empty_patch one-of patches with [any? turtles-here = false] move-to empty_patch ] set groupid 0 ] let unassigned turtles ;; start with group 1 and loop to build each group let current 1 while [any? unassigned] [ ;; place a randomly chosen set of group-size turtles into the current ;; group. or, if there are less than group-size turtles left, place the ;; rest of the turtles in the current group. ask n-of (min (list size_n (count unassigned))) unassigned [ set groupid current set n_neighbors other turtles with [groupid = current] ] ;; consider the next group. set current current + 1 ;; remove grouped turtles from the pool of turtles to assign set unassigned unassigned with [groupid = 0] ] ask turtles [ ;; if i'm in a group, move towards "home" for my group if groupid != 0 [ face get-home let p [neighbors] of get-home let area (patch-set get-home p) let my_patch one-of area move-to my_patch ] ;; wiggle a little and always move forward, to make sure turtles don't all ;; pile up lt random 5 rt random 5 fd 1 ] end ;; Courtesy of Uri Wilensky: ;; figures out the home patch for a group. this looks complicated, but the ;; idea is simple. we just want to lay the groups out in a regular grid, ;; evenly spaced throughout the world. we want the grid to be square, so in ;; some cases not all the positions are filled. to-report get-home ;; turtle procedure ;; calculate the minimum length of each side of our grid let side ceiling (sqrt (max [groupid] of turtles + 1)) report patch ;; compute the x coordinate (round ((world-width / side) * (groupid mod side) + min-pxcor + int (world-width / (side * 2)))) ;; compute the y coordinate (round ((world-height / side) * int (groupid / side) + min-pycor + int (world-height / (side * 2)))) end to PD_pairing ;; Pairwise Prisoner's Dilemma Game ask turtles [ let partner find-partner ;if partner = nobody [die] ;; dies if isolated! let utility 0 let total_cost 0 let total_benefit 0 let personal_cost 0 ifelse member? self cooperators [set personal_cost cost] [set personal_cost 0] set total_cost total_cost + personal_cost ifelse member? partner cooperators [set total_benefit total_benefit + benefit ;; if partner is a cooperator, add benefit to 'totalbenefit' recorder. set utility utility + Benefit - personal_cost] ;; if neighbor is a cooperator, then add benefit... [set utility utility - personal_cost] ;;if neighbor is a defector, then no benefit and subtract personal cost, if any... set payoff utility set earnings earnings + payoff set mycosts total_cost set mybenefits total_benefit ] end To Public_goods_game foreach groups [ ?1 -> let group_share 0 let thisgroup turtles with [groupid = ?1] ask thisgroup [ set mycosts 0 let t group_coop let r random-float 1 ;; ERROR ifelse contrite > 0 [ ;; if contrite > 0, then cooperate, unconditionally, otherwise... set breed cooperators set shape "face happy" set size 1 set color yellow ;; then cooperate set group_share group_share + Benefit set mycosts cost] ;;ERROR IS BOTH ERROR TOWARD COOPERATING AND ERROR TOWARD DEFECTING. [ifelse t >= t_threshold ;;if enough other group members contributed last round then COOPERATE. [ifelse r <= error_rate[ ;; HERE, ERROR MEANS DEFECTING INSTEAD OF COOPERATING set breed defectors set shape "face sad" set size 1.5 set color red if contrite? = true [if r <= error_rate AND t >= t_threshold [set contrite 2 ]] ] [set breed cooperators set shape "face happy" set size 1 set color yellow ;; then cooperate set group_share group_share + Benefit set mycosts cost]] [ifelse r <= error_rate[ ;; HERE, ERROR MEANS COOPERATING INSTEAD OF DEFECTING set breed cooperators set shape "face happy" set size 1 set color yellow set group_share group_share + Benefit set mycosts cost] [set breed defectors set shape "face sad" set size 1.5 set color red]]] ] ask thisgroup [ set payoff (group_share / (size_n - 1)) - mycosts ;; payoff is b/n or b/(n-1) ?? set earnings earnings + payoff set group_coop count cooperators with [groupid = ?1] ] ] end to replicator_dynamics if Replicator_options = "Relative Payoff" [Relative_Payoff] if Replicator_options = "Variance Ratio" [Variance_Replicator] if Replicator_options = "Replicator Equation" [Replicator_equation] if Replicator_options = "Imitation" [Imitate] end ;; probability of changing to another strategy is proportional to the difference between the *mean* payoffs for defectors and cooperators. ;; turtle only can switch if the payoffs are larger for the other strategy. to Relative_payoff ifelse mean [payoff] of cooperators > mean [payoff] of defectors [ ;; if cooperators making more payoff, then select the defectors to change ask defectors [let pr random-float 1 if pr <= RD1 [delete_defectors]]] [ ;; if defectors making more, then ask cooperators to change ask cooperators [let pr random-float 1 if pr <= RD1 [delete_cooperators]]] end to dying-turtles ;; turtles die if their earnings (or possibly their payoffs) get below zero. let consuming ((benefit - cost) / size_n) / 2 ask turtles [ set earnings earnings - consuming if earnings < 0 [die]] ;ask turtles [setup-neighbors] ;; must reset potential partners to avoid calling on dead turtles! end to kill-d ;;RULE This just means that half the cost is deducted from earnings each round a turtle has no cooperators to cooperate with let consuming ((benefit - cost) / size_n) / 2 ask turtles [ let g 0 ask N_neighbors [if member? self cooperators [set g g + 1]] if g = 0 [set earnings earnings - consuming] ] end to Variance_replicator ;; based on variable 'popchange' ;; According to Bowles and Gintis, the ratio of between-group variation (of altruists) to the total variation (which is the weighted-average within-group variation + the between- ;; group variation) must be greater than the ratio c/b for evolution to favor altruism. ;; This ratio is also the probability of being paired with an altruist minus the probability of being paired with an altruist conditional on being an altruist or non-altruist, ;; respectively, or P(A|A) - P(A|N). This seems more of a predictive tool than an algorithm to change the population. variances let c count turtles let new_agents pop_change * c ;; let c_r round new_agents ifelse c_r > 0 [;; add more cooperators, kill defectors let c_d count defectors let c_min min (list c_r c_d) let deleted_defectors min-n-of c_min defectors [payoff] ask deleted_defectors [delete_defectors]] ;;add more defectors, kill cooperators [let p_cr c_r * -1 ;; convert to a positive number let c_c count cooperators let c_min min (list p_cr c_c) let deleted_cooperators min-n-of c_min cooperators [payoff] ask deleted_cooperators [delete_cooperators] ] end to Replicator_equation ;; let Pr(i) = the proportion of strategy i ;; let $i = the payoff of strategy i, since I can't write the pi symbol here. ;; the new proportion of strategy i in the population at time t+1 is given by: ;; Pr(i)t+1 = Pr(i)$(i) / Sum of Weights ;; the weight for each strategy is given by the numerator let expected_coop_change coop_pay - (count cooperators / count turtles) let expected_defect_change coop_def - (count defectors / count turtles) let c expected_coop_change * count turtles ;; gives the number of turtles that will be changed let c_r round c ;rounded ifelse c_r > 0 [ ;; add more cooperators, kill defectors let c_d count defectors let c_min min (list c_r c_d) let deleted_defectors min-n-of c_min defectors [payoff] ask deleted_defectors [delete_defectors]] ;;add more defectors, kill cooperators [let p_cr c_r * -1 ;; convert to a positive number let c_c count cooperators let c_min min (list p_cr c_c) let deleted_cooperators min-n-of c_min cooperators [payoff] ask deleted_cooperators [delete_cooperators] ] end to imitate ;; this probably won't work, because its not clear how turtles will decide to imitate.. ;; if all agents imitate most successful agent in their group, then it creates immediate within-group homogeneity ;setting it initially to 4 closest agents, von Neuman, or Moore neighborhood, can't remember which. ask turtles [ let other_a min-n-of 4 other turtles [distance self] let max_a max-one-of other_a [payoff] if [payoff] of max_a > [payoff] of self [ ifelse [breed] of max_a = cooperators [delete_defectors] [delete_cooperators] set t_threshold [t_threshold] of max_a ;; copying the threshold (for public goods games), not just the strategy! set group_coop t_threshold ] ] end to delete_defectors ;; hatch and die let i [groupid] of self hatch-cooperators 1 [ set groupid i let mygroup other turtles with [groupid = i] ;create-wlinks-with mygroup let cnt size_n ;; (i.e. n) let t random cnt ;; t will be automatically between 0 and n and therefore not a defector set t_threshold t set color yellow set size .5 set group_coop t_threshold ;; will initially act as if just enough turtles have cooperated in previous round ] die end to delete_cooperators let i [groupid] of self hatch-defectors 1 [ set groupid i let mygroup other turtles with [groupid = i] ;create-wlinks-with mygroup set t_threshold size_n + 1 ;; requires more turtles to cooperate than actually exist, therefore a defector set shape "face sad" set size 1 set color red set group_coop 0 ;; will initially act as if just enough turtles have cooperated in previous round ] die end to variances let jmin min [groupid] of turtles let jmax max [groupid] of turtles let j jmin let avgrouplist [] let bgrouplist [] repeat jmax [ let grouplist [] ask turtles with [groupid = j] [ ifelse [breed] of self = cooperators [set grouplist fput 1 grouplist] [set grouplist fput 0 grouplist] ;; set 1 if altruist, 0 otherwise ] ask turtles with [groupid = j] [ set wingroupvar variance grouplist ] set j j + 1 ] let j2 min [groupid] of turtles repeat jmax [ let num count turtles with [groupid = j2] let numi count turtles with [groupid = j2 AND breed = cooperators] ;; counts number of cooperators let pj numi / num ;; frequency of altruists in the group let f num / count turtles let gvar mean [wingroupvar] of turtles with [groupid = j2] ;; every turtle in the group should have the same within group variance, but just in case, i take the average here. set avgrouplist fput (f * gvar) avgrouplist set bgrouplist fput pj bgrouplist set j2 j2 + 1 ] set avwithingroupvar variance avgrouplist ;; reports the weighted-average within-group variance of altruists set betweenvar variance bgrouplist set variance_ratio betweenvar / (avwithingroupvar + betweenvar) p_change end to p_change ;; change in the fraction of altruists population in total population let b Benefit let c Cost let var_pj betweenvar let var_pij avwithingroupvar let p ((b - c) * var_pj) - (c * var_pij) set pop_change p end to-report coop_pay;; proportion of cooperators*payoff of cooperators divided by sum of weighted payoffs let expected_coop (count cooperators / count turtles) * mean [payoff] of cooperators let expected_def (count defectors / count turtles) * mean [payoff] of defectors let total_payoff_c expected_coop / (expected_def + expected_coop) report total_payoff_c end to-report coop_def let expected_coop (count cooperators / count turtles) * mean [payoff] of cooperators let expected_def (count defectors / count turtles) * mean [payoff] of defectors let total_payoff_d expected_def / (expected_def + expected_coop) report total_payoff_d end to-report RD1 ;; veresion 3. Qij = B($j - $i) ;; probability that agent will switch from less profitable strategy to more profitable strategy ;; B has to be sufficiently small so that Qij is always <= 1 ! let B .1 ;; just trying random numbers let payoff_c mean [payoff] of cooperators let payoff_d mean [payoff] of defectors ifelse payoff_c > payoff_d [ ;; probability that defectors will switch to cooperation... let Qij B * (payoff_c - payoff_d) report Qij] [ ;; probability that cooperators will switch to defection... let Qij B * (payoff_d - payoff_c) report Qij] end to-report RD2 ;; Replicator Dynamics Version #2 for Cooperators ;; Pr(i)t+1 = Pr(i) - a * Pr(i)(1-P)B($j - $i) let B .1 ;; randomly assigned let p_c (count cooperators / count turtles) ;; proportion of turtles that are cooperators let p_d (count defectors / count turtles) ;; proportion defectors let payoff_c mean [payoff] of cooperators let payoff_d mean [payoff] of defectors let expected_p p_c - ( p_c * (1 - p_d) * B * (payoff_d - payoff_c)) report expected_p end
There are 2 versions of this model.
Attached files
File | Type | Description | Last updated | |
---|---|---|---|---|
A Cooperative Species.png | preview | Preview for 'A Cooperative Species' | about 7 years ago, by John Bradford | Download |
This model does not have any ancestors.
This model does not have any descendants.